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              <p>ABSTRACT: INTRODUCTION: Proteins encoded by Fanconi anemia (FA) and/or breast cancer (BrCa) susceptibility genes cooperate in a common DNA damage repair signaling pathway. To gain deeper insight into this pathway and its influence on cancer risk, we searched for novel components through protein physical interaction screens. METHODS: Protein physical interactions were screened using the yeast two-hybrid system. Co-affinity purifications and endogenous co-immunoprecipitation assays were performed to corroborate interactions. Biochemical and functional assays in human, mouse and Caenorhabditis elegans models were carried out to characterize pathway components. Thirteen FANCD2-monoubiquitinylation-positive FA cell lines excluded for genetic defects in the downstream pathway components and 300 familial BrCa patients negative for BRCA1/2 mutations were analyzed for genetic mutations. Common genetic variants were genotyped in 9,573 BRCA1/2 mutation carriers for associations with BrCa risk. RESULTS: A previously identified co-purifying protein with PALB2 was identified, MRG15 (MORF4L1 gene). Results in human, mouse and C. elegans models delineate molecular and functional relationships with BRCA2, PALB2, RAD51 and RPA1 that suggest a role for MRG15 in the repair of DNA double-strand breaks. Mrg15-deficient murine embryonic fibroblasts showed moderate sensitivity to γ-irradiation relative to controls and reduced formation of Rad51 nuclear foci. Examination of mutants of MRG15 and BRCA2 C. elegans orthologs revealed phenocopy by accumulation of RPA-1 (human RPA1) nuclear foci and aberrant chromosomal compactions in meiotic cells. However, no alterations or mutations were identified for MRG15/MORF4L1 in unclassified FA patients and BrCa familial cases. Finally, no significant associations between common MORF4L1 variants and BrCa risk for BRCA1 or BRCA2 mutation carriers were identified: rs7164529, Ptrend = 0.45 and 0.05, P2df = 0.51 and 0.14, respectively; and rs10519219, Ptrend = 0.92 and 0.72, P2df = 0.76 and 0.07, respectively. CONCLUSIONS: While the present study expands on the role of MRG15 in the control of genomic stability, weak associations cannot be ruled out for potential low-penetrance variants at MORF4L1 and BrCa risk among BRCA2 mutation carriers.</p>
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        </org>
        <org type="institution" xml:id="struct-342726" status="INCOMING">
          <orgName>Central Manchester University Hospitals</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342594" status="INCOMING">
          <orgName>The Institute of Cancer Research and Royal Marsden NHS Foundation Trust</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-575091" status="VALID">
          <orgName>Guy's and St Thomas' NHS Foundation Trust</orgName>
          <desc>
            <address>
              <addrLine>St Thomas' Hospital, Westminster Bridge Rd, London SE1 7EH, Royaume-Uni</addrLine>
              <country key="GB"/>
            </address>
            <ref type="url">https://www.guysandstthomas.nhs.uk/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-308690" status="INCOMING">
          <orgName>St James's hospital</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342727" status="INCOMING">
          <orgName>Ferguson-Smith Centre for Clinical Genetics</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342596" status="VALID">
          <orgName>Birmingham Women's and Children's NHS Foundation Trust</orgName>
          <desc>
            <address>
              <addrLine>Birmingham Women's HospitalMindelsohn WayBirmingham B15 2TG Birmingham Children's HospitalSteelhouse LaneBirmingham B4 6NH</addrLine>
              <country key="GB"/>
            </address>
            <ref type="url">https://bwc.nhs.uk/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-341095" status="VALID">
          <idno type="ROR">https://ror.org/02md8hv62</idno>
          <orgName>Sheffield Children's NHS Foundation Trust</orgName>
          <desc>
            <address>
              <addrLine>Sheffield</addrLine>
              <country key="GB"/>
            </address>
            <ref type="url">https://www.sheffieldchildrens.nhs.uk/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342760" status="VALID">
          <idno type="ROR">https://ror.org/05p40t847</idno>
          <orgName>Newcastle Upon Tyne Hospitals NHS Foundation Trust</orgName>
          <desc>
            <address>
              <addrLine>Queen Victoria Road, Newcastle upon Tyne, NE1 4LP</addrLine>
              <country key="GB"/>
            </address>
            <ref type="url">http://www.newcastle-hospitals.org.uk/index.aspx</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-303063" status="VALID">
          <idno type="ROR">https://ror.org/026zzn846</idno>
          <orgName>Queen Mary University of London</orgName>
          <orgName type="acronym">QMUL</orgName>
          <desc>
            <address>
              <addrLine>Mile End Road, London E1 4NS</addrLine>
              <country key="GB"/>
            </address>
            <ref type="url">http://www.qmul.ac.uk/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-308787" status="INCOMING">
          <orgName>St George's Hospital</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342602" status="INCOMING">
          <orgName>Royal Devon &amp; Exeter Hospital</orgName>
          <desc>
            <address>
              <country key="GB"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-333925" status="VALID">
          <idno type="ROR">https://ror.org/02405mj67</idno>
          <orgName>Belfast City Hospital</orgName>
          <desc>
            <address>
              <addrLine>Lisburn Rd, Belfast BT9 7AB, Royaume-Uni</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">https://belfasttrust.hscni.net/hospitals/bch/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-335184" status="VALID">
          <idno type="ISNI">0000 0004 0624 9907</idno>
          <idno type="ROR">https://ror.org/009kr6r15</idno>
          <idno type="Wikidata">Q7987867</idno>
          <orgName>Western General Hospital</orgName>
          <date type="start">1868-01-01</date>
          <desc>
            <address>
              <addrLine>Crewe Rd S, Edinburgh EH4 2XU</addrLine>
              <country key="GB"/>
            </address>
            <ref type="url">https://www.nhslothian.scot/goingtohospital/western-general-hospital/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342603" status="INCOMING">
          <orgName>Department of Preventive and Predictive Medicine</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342761" status="INCOMING">
          <orgName>Fondazione IRCCS Istituto Nazionale Tumori (INT)</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-117093" status="VALID">
          <orgName>IFOM, Istituto FIRC di Oncologia Molecolare</orgName>
          <orgName type="acronym">IFOM</orgName>
          <desc>
            <address>
              <addrLine>Via Adamello 16, 20139 - Milan, Italy</addrLine>
              <country key="IT"/>
            </address>
            <ref type="url">https://www.ifom.eu/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342762" status="INCOMING">
          <orgName>Fondazione IRCCS INT</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342605" status="INCOMING">
          <orgName>Istituto Europeo di Oncologia (IEO)</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-380289" status="VALID">
          <orgName>Istituto di Ricovero e Cura a Carattere Scientifico</orgName>
          <orgName type="acronym">IRCCS</orgName>
          <desc>
            <address>
              <addrLine>Via Conte Ruggiero, 73, 94018 Troina EN</addrLine>
              <country key="IT"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-47709" status="VALID">
          <idno type="IdRef">027548325</idno>
          <idno type="ROR">https://ror.org/048tbm396</idno>
          <orgName>Università degli studi di Torino = University of Turin</orgName>
          <orgName type="acronym">UNITO</orgName>
          <date type="start">1404-01-01</date>
          <desc>
            <address>
              <addrLine>Via Verdi, 8 10124 Torino Italie</addrLine>
              <country key="IT"/>
            </address>
            <ref type="url">http://www.unito.it/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-229915" status="VALID">
          <idno type="IdRef">026430711</idno>
          <idno type="ROR">https://ror.org/02be6w209</idno>
          <orgName>Università degli Studi di Roma "La Sapienza" = Sapienza University [Rome]</orgName>
          <orgName type="acronym">UNIROMA</orgName>
          <date type="start">1303-01-01</date>
          <desc>
            <address>
              <addrLine>Piazzale Aldo Moro 5, 00185 Roma</addrLine>
              <country key="IT"/>
            </address>
            <ref type="url">https://www.uniroma1.it/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-148408" status="VALID">
          <idno type="IdRef">026430584</idno>
          <idno type="ROR">https://ror.org/04jr1s763</idno>
          <orgName>Università degli Studi di Firenze = University of Florence = Université de Florence</orgName>
          <orgName type="acronym">UniFI</orgName>
          <desc>
            <address>
              <addrLine>Piazza di San Marco, 4, 50121 Firenze FI, Italie</addrLine>
              <country key="IT"/>
            </address>
            <ref type="url">https://www.unifi.it/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-327359" status="INCOMING">
          <orgName>Fiorgen Foundation for Pharmacogenomics</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-330035" status="INCOMING">
          <orgName>Regina Elena Cancer Institute</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="regrouplaboratory" xml:id="struct-441517" status="VALID">
          <orgName>Instituto Español de Oceanografía = Spanish Institute of Oceanography</orgName>
          <orgName type="acronym">IEO | CSIC</orgName>
          <desc>
            <address>
              <addrLine>Consejo Superior de Investigaciones Científicas | C. del Corazón de María, 8, Chamartín, 28002 Madrid</addrLine>
              <country key="ES"/>
            </address>
            <ref type="url">https://www.ieo.es/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-106380" type="direct"/>
          </listRelation>
        </org>
        <org type="regroupinstitution" xml:id="struct-106380" status="VALID">
          <idno type="IdRef">026373246</idno>
          <idno type="ISNI">0000000123535612</idno>
          <idno type="ROR">https://ror.org/02gfc7t72</idno>
          <idno type="Wikidata">Q74202089</idno>
          <orgName>Consejo Superior de Investigaciones Cientificas [España] = Spanish National Research Council [Spain]</orgName>
          <orgName type="acronym">CSIC</orgName>
          <date type="start">1939-01-01</date>
          <desc>
            <address>
              <addrLine>Serrano 117, 28006, Madrid</addrLine>
              <country key="ES"/>
            </address>
            <ref type="url">http://www.csic.es/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-140553" status="VALID">
          <idno type="ROR">https://ror.org/004y8wk30</idno>
          <orgName>Queensland Institute of Medical Research</orgName>
          <desc>
            <address>
              <addrLine>Queensland Institute of Medical Research Locked Bag 2000 Royal Brisbane Hospital Herston, Qld 4029 Australia</addrLine>
              <country key="AU"/>
            </address>
            <ref type="url">http://www.qimr.edu.au/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-307782" status="INCOMING">
          <orgName>The Netherlands Cancer Institute</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-308157" status="VALID">
          <idno type="ROR">https://ror.org/018906e22</idno>
          <orgName>Erasmus University Medical Center [Rotterdam]</orgName>
          <orgName type="acronym">Erasmus MC</orgName>
          <desc>
            <address>
              <addrLine>Rotterdam</addrLine>
              <country key="NL"/>
            </address>
            <ref type="url">http://www.erasmusmc.nl</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342591" status="INCOMING">
          <orgName>Family Cancer Clinic</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-300851" status="VALID">
          <idno type="ROR">https://ror.org/05xvt9f17</idno>
          <orgName>Leiden University Medical Center [Leiden]</orgName>
          <orgName type="acronym">LUMC</orgName>
          <desc>
            <address>
              <addrLine>Leids Universitair Medisch Centrum, Universiteit Leiden - 1233 ZA, Leiden</addrLine>
              <country key="NL"/>
            </address>
            <ref type="url">https://www.lumc.nl</ref>
          </desc>
          <listRelation>
            <relation active="#struct-121723" type="direct"/>
          </listRelation>
        </org>
        <org type="regroupinstitution" xml:id="struct-121723" status="VALID">
          <idno type="ROR">https://ror.org/027bh9e22</idno>
          <orgName>Universiteit Leiden = Leiden University</orgName>
          <desc>
            <address>
              <addrLine>Leiden University | 2300 RA Leiden The Netherlands</addrLine>
              <country key="NL"/>
            </address>
            <ref type="url">https://www.universiteitleiden.nl/en/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-305057" status="VALID">
          <idno type="ROR">https://ror.org/05wg1m734</idno>
          <orgName>Radboud University Medical Center [Nijmegen]</orgName>
          <orgName type="acronym">RadboudUMC</orgName>
          <desc>
            <address>
              <addrLine>PO Box 9101, 6500 HB Nijmegen, The Netherlands</addrLine>
              <country key="NL"/>
            </address>
            <ref type="url">https://www.radboudumc.nl/EN</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-73956" status="VALID">
          <idno type="ROR">https://ror.org/0575yy874</idno>
          <orgName>University Medical Center [Utrecht]</orgName>
          <orgName type="acronym">UMCU</orgName>
          <desc>
            <address>
              <addrLine>Heidelberglaan 1003584 CX Utrecht</addrLine>
              <country key="NL"/>
            </address>
            <ref type="url">https://www.umcutrecht.nl/en/</ref>
          </desc>
        </org>
        <org type="regrouplaboratory" xml:id="struct-112034" status="VALID">
          <orgName>Academic Medical Center - Academisch Medisch Centrum [Amsterdam]</orgName>
          <orgName type="acronym">AMC</orgName>
          <desc>
            <address>
              <addrLine>Meibergdreef 9 1105 AZ Amsterdam</addrLine>
              <country key="NL"/>
            </address>
            <ref type="url">https://www.amc.nl/web/home.htm</ref>
          </desc>
          <listRelation>
            <relation active="#struct-120654" type="direct"/>
          </listRelation>
        </org>
        <org type="regroupinstitution" xml:id="struct-120654" status="VALID">
          <idno type="IdRef">026431610</idno>
          <idno type="ISNI">0000 0000 8499 2262</idno>
          <idno type="ROR">https://ror.org/04dkp9463</idno>
          <idno type="Wikidata">Q214341</idno>
          <orgName>Universiteit van Amsterdam = University of Amsterdam</orgName>
          <orgName type="acronym">UvA</orgName>
          <date type="start">1632-01-01</date>
          <desc>
            <address>
              <addrLine>Spui 21 1012 WX Amsterdam</addrLine>
              <country key="NL"/>
            </address>
            <ref type="url">https://www.uva.nl/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-344060" status="VALID">
          <orgName>Maastricht University Medical Center+</orgName>
          <orgName type="acronym">Maastricht UMC+</orgName>
          <desc>
            <address>
              <country key="NL"/>
            </address>
            <ref type="url">https://www.mumc.nl/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-327362" status="INCOMING">
          <orgName>VU Medical Center</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="regrouplaboratory" xml:id="struct-329105" status="VALID">
          <orgName>School of Medicine [University of Utah, Salt Lake City]</orgName>
          <desc>
            <address>
              <addrLine>30 N. 1900 E.Salt Lake City, UT 84132</addrLine>
              <country key="US"/>
            </address>
            <ref type="url">http://medicine.utah.edu/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-300863" type="direct"/>
          </listRelation>
        </org>
        <org type="institution" xml:id="struct-300863" status="VALID">
          <idno type="ROR">https://ror.org/03r0ha626</idno>
          <orgName>University of Utah</orgName>
          <desc>
            <address>
              <addrLine>201 Presidents Cir, Salt Lake City, UT 84112</addrLine>
              <country key="US"/>
            </address>
            <ref type="url">http://www.utah.edu/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-313617" status="INCOMING">
          <orgName>Huntsman Cancer Institute</orgName>
          <desc>
            <address>
              <addrLine>Salk Lake City</addrLine>
              <country key="US"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342763" status="INCOMING">
          <orgName>Cancer Prevention Institute of California</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-142307" status="VALID">
          <idno type="ROR">https://ror.org/02jzgtq86</idno>
          <orgName>Dana-Farber Cancer Institute [Boston]</orgName>
          <desc>
            <address>
              <addrLine>450 Brookline Ave.Boston, MA 02215</addrLine>
              <country key="US"/>
            </address>
            <ref type="url">http://www.dana-farber.org/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-130467" status="VALID">
          <idno type="ROR">https://ror.org/03wevmz92</idno>
          <orgName>Harvard Medical School [Boston]</orgName>
          <orgName type="acronym">HMS</orgName>
          <desc>
            <address>
              <addrLine>25 Shattuck Street Boston, MA 02115</addrLine>
              <country key="US"/>
            </address>
            <ref type="url">http://www.hms.harvard.edu</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-306322" status="VALID">
          <idno type="ROR">https://ror.org/01ej9dk98</idno>
          <orgName>University of Melbourne</orgName>
          <desc>
            <address>
              <addrLine>Parkville VIC 3010</addrLine>
              <country key="AU"/>
            </address>
            <ref type="url">http://www.unimelb.edu.au/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342764" status="INCOMING">
          <orgName>Melbourne School of Population Health</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-310911" status="INCOMING">
          <orgName>Fox Chase Cancer Center</orgName>
          <desc>
            <address>
              <country key="US"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342765" status="INCOMING">
          <orgName>Skåne University Hospital</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-344927" status="VALID">
          <idno type="ROR">https://ror.org/02z31g829</idno>
          <orgName>Skane University Hospital [Lund]</orgName>
          <desc>
            <address>
              <addrLine>Getingevägen 4, 222 41 Lund</addrLine>
              <country key="SE"/>
            </address>
            <ref type="url">https://www.skane.se/sv/Webbplatser/Skanes-universitetssjukhus/Skane-University-Hospital/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-307368" status="VALID">
          <idno type="ROR">https://ror.org/00m8d6786</idno>
          <orgName>Karolinska University Hospital [Stockholm]</orgName>
          <desc>
            <address>
              <addrLine>Huddinge141 86 Stockholm</addrLine>
              <country key="SE"/>
            </address>
            <ref type="url">http://karolinska.se/karolinska-university-hospital/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-50873" status="VALID">
          <idno type="IdRef">026431661</idno>
          <idno type="ROR">https://ror.org/048a87296</idno>
          <orgName>Uppsala University</orgName>
          <desc>
            <address>
              <addrLine>Box 256, SE-751 05 Uppsala</addrLine>
              <country key="SE"/>
            </address>
            <ref type="url">http://www.uu.se/en/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-306574" status="INCOMING">
          <orgName>University Hospital</orgName>
          <desc>
            <address>
              <addrLine>Linköping</addrLine>
              <country key="SE"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342766" status="INCOMING">
          <orgName>Hälsouniversitetet Universitetssjukhuset</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-329653" status="VALID">
          <orgName>Chaim Sheba Medical Center</orgName>
          <desc>
            <address>
              <country key="IL"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-329290" status="INCOMING">
          <orgName>Institute of Human Genetics</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="regroupinstitution" xml:id="struct-63133" status="VALID">
          <idno type="ROR">https://ror.org/04mhzgx49</idno>
          <orgName>Tel Aviv University</orgName>
          <orgName type="acronym">TAU</orgName>
          <desc>
            <address>
              <addrLine>Tel Aviv 699780</addrLine>
              <country key="IL"/>
            </address>
            <ref type="url">https://english.tau.ac.il/</ref>
          </desc>
        </org>
        <org type="regrouplaboratory" xml:id="struct-346739" status="VALID">
          <orgName>Perelman School of Medicine</orgName>
          <desc>
            <address>
              <addrLine>Philadelphia</addrLine>
              <country key="US"/>
            </address>
            <ref type="url">https://www.med.upenn.edu/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-115889" type="direct"/>
          </listRelation>
        </org>
        <org type="institution" xml:id="struct-115889" status="VALID">
          <idno type="ROR">https://ror.org/00b30xv10</idno>
          <orgName>University of Pennsylvania [Philadelphia]</orgName>
          <desc>
            <address>
              <addrLine>3451 Walnut Street, Philadelphia, PA 19104 | 215-898-5000</addrLine>
              <country key="US"/>
            </address>
            <ref type="url">http://www.upenn.edu/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342767" status="INCOMING">
          <orgName>Abramson Cancer Center</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-300837" status="VALID">
          <orgName>University of Iceland [Reykjavik]</orgName>
          <desc>
            <address>
              <addrLine>Sæmundargata 2, 101 Reykjavík</addrLine>
              <country key="IS"/>
            </address>
            <ref type="url">http://english.hi.is/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-310910" status="VALID">
          <orgName>Mayo Clinic</orgName>
          <desc>
            <address>
              <country key="US"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-166039" status="VALID">
          <orgName>Universität zu Lübeck = University of Luebeck [Lübeck]</orgName>
          <desc>
            <address>
              <addrLine>Ratzeburger Allee 160, 23562 Lübeck</addrLine>
              <country key="DE"/>
            </address>
            <ref type="url">https://www.uni-luebeck.de/en/university/university.html</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342607" status="INCOMING">
          <orgName>Center for Structural and Cell Biology in Medicine</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-326928" status="INCOMING">
          <orgName>Hospital Clínico San Carlos</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-301865" status="VALID">
          <orgName>CIBER de Enfermedades Raras (CIBERER)</orgName>
          <desc>
            <address>
              <country key="ES"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-323086" status="INCOMING">
          <orgName>Spanish National Cancer Research Centre</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-1082138" status="VALID">
          <orgName>Centro de Investigaciones Energéticas, Medioambientales y Tecnológicas</orgName>
          <orgName type="acronym">CIEMAT</orgName>
          <date type="start">1986-01-01</date>
          <desc>
            <address>
              <addrLine>Av. Complutense, 40, 28040 Madrid, Espagne</addrLine>
              <country key="ES"/>
            </address>
            <ref type="url">https://www.ciemat.es/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-534438" status="VALID">
          <idno type="ROR">https://ror.org/02e8hzf44</idno>
          <orgName>Helsinki University Hospital [Finland]</orgName>
          <orgName type="acronym">HUS</orgName>
          <desc>
            <address>
              <addrLine>Topeliuksenkatu 5, 00260 Helsinki.</addrLine>
              <country key="FI"/>
            </address>
            <ref type="url">https://www.hus.fi/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-150185" status="VALID">
          <orgName>German Cancer Research Center - Deutsches Krebsforschungszentrum [Heidelberg]</orgName>
          <orgName type="acronym">DKFZ</orgName>
          <desc>
            <address>
              <addrLine>German Cancer Research Center Im Neuenheimer Feld 280 D-69120 Heidelberg</addrLine>
              <country key="DE"/>
            </address>
            <ref type="url">https://www.dkfz.de/en/index.html</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-51187" status="VALID">
          <idno type="ROR">https://ror.org/03etyjw28</idno>
          <orgName>Pontificia Universidad Javeriana</orgName>
          <orgName type="acronym">PUJ</orgName>
          <date type="start">1937-01-01</date>
          <desc>
            <address>
              <addrLine>Bogotá</addrLine>
              <country key="CO"/>
            </address>
            <ref type="url">https://www.javeriana.edu.co/inicio</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-366408" status="VALID">
          <idno type="IdRef">026619385</idno>
          <idno type="ISNI">0000000417573729</idno>
          <idno type="ROR">https://ror.org/03ad39j10</idno>
          <idno type="Wikidata">Q645663</idno>
          <orgName>University of Pisa [Italy] = Università di Pisa [Italia] = Université de Pise [Italie]</orgName>
          <orgName type="acronym">UniPi</orgName>
          <date type="start">1343-01-01</date>
          <desc>
            <address>
              <addrLine>Lungarno Pacinotti 43 – 56126 Pisa – Italia</addrLine>
              <country key="IT"/>
            </address>
            <ref type="url">https://www.unipi.it/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-408606" status="VALID">
          <orgName>University of Kansas Medical Center [Kansas City, KS, USA]</orgName>
          <desc>
            <address>
              <addrLine>3901 Rainbow Boulevard, Kansas City, KS 66160</addrLine>
              <country key="US"/>
            </address>
            <ref type="url">http://www.kumc.edu/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342608" status="INCOMING">
          <orgName>N.N. Petrov Institute of Oncology</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-303686" status="VALID">
          <idno type="IdRef">027294560</idno>
          <idno type="ISNI">0000 0001 2160 3183</idno>
          <idno type="ROR">https://ror.org/03nadee84</idno>
          <orgName>Vilnius University [Vilnius]</orgName>
          <date type="start">1579-01-01</date>
          <desc>
            <address>
              <addrLine>3 Universiteto St, LT-01513 Vilnius</addrLine>
              <country key="LT"/>
            </address>
            <ref type="url">http://www.vu.lt/en/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-342768" status="INCOMING">
          <orgName>Hospital Santariskiu Clinics</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-20549" status="VALID">
          <idno type="ROR">https://ror.org/01cmnjq37</idno>
          <orgName>Centre Léon Bérard [Lyon]</orgName>
          <desc>
            <address>
              <addrLine>28, rue Laennec 69373 LYON Cedex 08</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.centreleonberard.fr/</ref>
          </desc>
        </org>
        <org type="regroupinstitution" xml:id="struct-300074" status="VALID">
          <idno type="IdRef">026534657</idno>
          <idno type="ROR">https://ror.org/01502ca60</idno>
          <orgName>Hospices Civils de Lyon</orgName>
          <orgName type="acronym">HCL</orgName>
          <desc>
            <address>
              <addrLine>3 Quai des Célestins69002 Lyon</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.chu-lyon.fr/fr/hospices-civils-de-lyon-hcl</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-194495" status="VALID">
          <idno type="IdRef">026402823</idno>
          <idno type="ISNI">0000000121686185</idno>
          <idno type="ROR">https://ror.org/029brtt94</idno>
          <orgName>Université Claude Bernard Lyon 1</orgName>
          <orgName type="acronym">UCBL</orgName>
          <desc>
            <address>
              <addrLine>43, boulevard du 11 novembre 1918, 69622 Villeurbanne cedex</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.univ-lyon1.fr/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-301088" type="direct"/>
          </listRelation>
        </org>
        <org type="regroupinstitution" xml:id="struct-301088" status="VALID">
          <idno type="ROR">https://ror.org/01rk35k63</idno>
          <orgName>Université de Lyon</orgName>
          <desc>
            <address>
              <addrLine>92 rue Pasteur - CS 30122, 69361 Lyon Cedex 07</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">https://www.universite-lyon.fr/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-303623" status="VALID">
          <idno type="IdRef">026388278</idno>
          <idno type="ROR">https://ror.org/02vjkv261</idno>
          <orgName>Institut National de la Santé et de la Recherche Médicale</orgName>
          <orgName type="acronym">INSERM</orgName>
          <desc>
            <address>
              <addrLine>101, rue de Tolbiac, 75013 Paris</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.inserm.fr</ref>
          </desc>
        </org>
        <org type="regroupinstitution" xml:id="struct-441569" status="VALID">
          <idno type="IdRef">02636817X</idno>
          <idno type="ISNI">0000000122597504</idno>
          <idno type="ROR">https://ror.org/02feahw73</idno>
          <orgName>Centre National de la Recherche Scientifique</orgName>
          <orgName type="acronym">CNRS</orgName>
          <date type="start">1939-10-19</date>
          <desc>
            <address>
              <country key="FR"/>
            </address>
            <ref type="url">https://www.cnrs.fr/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-301888" status="VALID">
          <idno type="ISNI">0000 0004 0639 6384</idno>
          <idno type="ROR">https://ror.org/04t0gwh46</idno>
          <orgName>Institut Curie [Paris]</orgName>
          <desc>
            <address>
              <addrLine>26 rue d'Ulm - 75248 Paris</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">https://science.curie.fr/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-301664" status="OLD">
          <idno type="IdRef">026404788</idno>
          <idno type="ISNI">0000 0001 2188 0914 </idno>
          <idno type="ROR">https://ror.org/0220k9r37</idno>
          <orgName>Université Paris Descartes - Paris 5</orgName>
          <orgName type="acronym">UPD5</orgName>
          <date type="start">1971-01-01</date>
          <date type="end">2019-12-31</date>
          <desc>
            <address>
              <addrLine>12, rue de l'École de Médecine - 75270 Paris cedex 06</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.parisdescartes.fr/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-30615" status="VALID">
          <idno type="ISNI">0000 0004 1795 1689</idno>
          <idno type="ROR">https://ror.org/02pwnhd33</idno>
          <orgName>Centre Jean Perrin [Clermont-Ferrand]</orgName>
          <orgName type="acronym">UNICANCER/CJP</orgName>
          <desc>
            <address>
              <addrLine>58 rue Montalembert, 63011 Clermont-Ferrand</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.cjp.fr/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-502406" type="direct"/>
          </listRelation>
        </org>
        <org type="regroupinstitution" xml:id="struct-502406" status="VALID">
          <orgName>UNICANCER</orgName>
          <desc>
            <address>
              <addrLine>101, rue de Tolbiac, 75654 Paris Cedex 13</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.unicancer.fr/</ref>
          </desc>
        </org>
        <org type="regrouplaboratory" xml:id="struct-301886" status="VALID">
          <idno type="ISNI">0000 0001 0131 6312</idno>
          <idno type="ROR">https://ror.org/03xfq7a50</idno>
          <orgName>Centre Régional de Lutte contre le Cancer Oscar Lambret [Lille]</orgName>
          <orgName type="acronym">UNICANCER/Lille</orgName>
          <desc>
            <address>
              <addrLine>3, rue Frédéric Combemale, 59020 Lille</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">https://www.centreoscarlambret.fr/</ref>
          </desc>
          <listRelation>
            <relation active="#struct-374570" type="direct"/>
            <relation active="#struct-502406" type="direct"/>
          </listRelation>
        </org>
        <org type="regroupinstitution" xml:id="struct-374570" status="VALID">
          <idno type="IdRef">223446556</idno>
          <idno type="ISNI">0000 0001 2242 6780</idno>
          <idno type="ROR">https://ror.org/02kzqn938</idno>
          <idno type="Wikidata">Q3551621</idno>
          <orgName>Université de Lille</orgName>
          <desc>
            <address>
              <addrLine>EPE Université de Lille. -- 42 rue Paul Duez, 59000 Lille</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">https://www.univ-lille.fr/</ref>
          </desc>
        </org>
        <org type="institution" xml:id="struct-331179" status="INCOMING">
          <orgName>CH Chambéry</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-300165" status="VALID">
          <orgName>Centre Hospitalier Régional Universitaire de Besançon</orgName>
          <orgName type="acronym">CHRU Besançon</orgName>
          <desc>
            <address>
              <addrLine>3, boulevard A. Fleming - 25030 Besançon Cedex</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">http://www.chu-besancon.fr/</ref>
          </desc>
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        <org type="institution" xml:id="struct-305160" status="INCOMING">
          <orgName>Hôpital Saint-Jacques</orgName>
          <desc>
            <address>
              <country key="FR"/>
            </address>
          </desc>
        </org>
        <org type="institution" xml:id="struct-179009" status="VALID">
          <idno type="ROR">https://ror.org/00jpq0w62</idno>
          <orgName>Centre Hospitalier Régional Universitaire de Tours</orgName>
          <orgName type="acronym">CHRU Tours</orgName>
          <desc>
            <address>
              <addrLine>Centre Hospitalier Régional Universitaire de Tours2 Boulevard Tonnellé 37044 Tours cedex 9</addrLine>
              <country key="FR"/>
            </address>
            <ref type="url">https://www.chu-tours.fr/</ref>
          </desc>
        </org>
      </listOrg>
    </back>
  </text>
</TEI>